Biology · Reference sheet

Classification and Taxonomy

The three domains of life and the taxonomic hierarchy from domain to species, binomial nomenclature, and how shared characteristics group organisms together.

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Why classify living things?

Earth holds millions of species, so biologists need an organized system to name them, group them, and study how they relate to one another. Taxonomy is the science of classifying living things into a nested hierarchy of ranks, based mostly on shared characteristics and shared ancestry. A clear classification system lets a scientist in Brazil and a scientist in Japan know they are studying the exact same organism, even if their everyday languages have no word in common.

Founder of modern taxonomy

The Swedish scientist Carl Linnaeus developed the classification system biologists still build on today, in the 1700s. His two big contributions were the nested hierarchy of ranks and the two-part naming system, both covered in this sheet.

The three domains

The domain is the broadest, most general rank in classification. Every organism on Earth belongs to one of three domains, sorted by cell type and basic biochemistry. Bacteria and Archaea are both made of prokaryotic cells, cells with no nucleus and no membrane-bound organelles, but they differ sharply in cell wall chemistry, membrane lipids, and genetics. Eukarya contains every organism built from eukaryotic cells, cells that do have a nucleus and membrane-bound organelles.

DomainCell typeNucleus?Example organisms
BacteriaProkaryoticNoEscherichia coli, Streptococcus
ArchaeaProkaryoticNoMethanogens, salt-loving and heat-loving extremophiles
EukaryaEukaryoticYesAnimals, plants, fungi, protists

Archaea aren't just bacteria that live in extreme places

Archaea were first noticed in extreme environments like hot springs and salt lakes, which is where the domain gets its reputation. But Archaea also live in ordinary places, including soil and the human gut. What actually separates Archaea from Bacteria is their cell wall and membrane chemistry, not where they happen to live.

The taxonomic hierarchy

Within Eukarya, and within each domain generally, organisms are sorted through a nested series of ranks, from the broadest group down to the most specific. Each rank sits fully inside the one above it. Every organism in a Genus shares that same Family, Order, Class, Phylum, Kingdom, and Domain, but a Genus can hold several different Species.

RankWhat it groupsBroad or specific
DomainThe most basic cell type and biochemistryBroadest
KingdomMajor body plan and lifestyle group
PhylumBody plan shared across many classes
ClassA major branch within a phylum
OrderA group of related families
FamilyA group of closely related genera
GenusA small group of very closely related species
SpeciesOrganisms that can interbreed and produce fertile offspringMost specific

A memory trick for the order

"Dear King Philip Came Over For Good Soup" gives the first letter of each rank in order: Domain, Kingdom, Phylum, Class, Order, Family, Genus, Species. Any sentence with eight words that start with those letters, in that order, works just as well.

Binomial nomenclature

Binomial nomenclature is the two-part scientific naming system Linnaeus introduced, and every recognized species has exactly one. The name always has two parts: the genus name first, Capitalized, and the species name second, lowercase. In print, scientific names are italicized. Once a name has been written in full once in a piece of writing, later mentions can abbreviate the genus to a single capital letter followed by a period. Example: The gray wolf's scientific name is *Canis lupus*. Genus Canis, species lupus. After the first full mention, it can be shortened to *C. lupus*. Example: The bacterium *Escherichia coli* is almost always abbreviated *E. coli* after its first mention, even in casual writing, because the full name is long and the organism comes up so often.

Common names cause confusion, scientific names don't

A single organism can have several common names depending on the region, and one common name can refer to more than one species. "Corn" means one crop in the United States and a different one in the United Kingdom, where it can mean wheat. The scientific name always points to exactly one species, no matter who is reading it or what language they speak.

How organisms get grouped together

Taxonomists group organisms by shared characteristics, and today those characteristics fall into two main kinds. Anatomical evidence compares body structures. Homologous structures are body parts that share the same underlying structure and the same evolutionary origin, even if they now do different jobs, such as a human arm, a bat wing, and a whale flipper, which all share the same basic bone arrangement inherited from a common ancestor. Molecular evidence compares DNA and protein sequences directly. Two species with more similar DNA sequences share a more recent common ancestor, and modern taxonomy leans heavily on this kind of comparison to confirm or correct groupings that were first made from anatomy alone.

Structure comparisonSame origin?What it tells you
Human arm and bat wing (homologous)YesBoth inherited from a shared ancestor, evidence of common ancestry
Bird wing and insect wing (analogous)NoSimilar function evolved independently, not evidence of common ancestry

Closer ranks mean closer relatives

Two species placed in the same Genus share a more recent common ancestor than two species that only share the same Phylum. The fewer ranks of separation between two organisms, the more recently their lineages split apart.

Following an organism through the hierarchy

Here is the full classification of the gray wolf, *Canis lupus*, read from the broadest rank down to the most specific.

RankClassification
DomainEukarya
KingdomAnimalia
PhylumChordata
ClassMammalia
OrderCarnivora
FamilyCanidae
GenusCanis
Specieslupus

Example: Compare the gray wolf (*Canis lupus*) to the domestic dog (*Canis familiaris*). They share the same Domain, Kingdom, Phylum, Class, Order, Family, and Genus, and only split apart at Species, which shows they are extremely close relatives. Now compare the gray wolf to a human (*Homo sapiens*). Both share Domain Eukarya, Kingdom Animalia, Phylum Chordata, and Class Mammalia, but they split apart at Order, wolves in Carnivora and humans in Primates. That earlier split tells you humans and wolves share a more distant common ancestor than a wolf and a dog do.

Try it yourself

A dichotomous key for identifying 8 fictional field specimens, one step at a time.

The trait values for the 8 specimens the key identifies.

Open the files in two windows

Open dichotomous-key.txt in any text editor and specimen-traits.csv in Google Sheets or Excel side by side. Start at Step 1 of the key, find the specimen's value for the named trait in the spreadsheet, and follow whichever line matches until you reach a name instead of a "go to step" instruction.

Using dichotomous-key.txt and specimen-traits.csv: 1. Walk specimen 1 through the key from Step 1. What is its identification? 2. Walk specimen 4 through the key from Step 1. What is its identification? 3. Specimens 6 and 7 both have no wings, live on land, and don't have 8 legs, so both reach Step 6. Which trait and value separates them there, and what is each one's identification? 4. Follow the key until it names a specimen "Rockshell." Which specimen number is it, and what two trait values (in the order the key checks them) led there? 5. Two specimens have wings. Identify both of them, and state the single trait (and its two different values) that tells them apart at Step 2.